Answer: columella

AnswerCrossword Clue
columellaThe axis of the spore-producing body of some lower plants
columellaThe pillar around which the cochlea spirals
columellaA structure resembling a small column and typically forming a central axis; in particular
columellaAn ossicle of the middle ear of birds, reptiles, and amphibians
columella(Latin) the uvula
columellaThe axis of a spiral shell