Answer: rhachis

AnswerCrossword Clue
rhachisthe part of the shaft of a feather bearing the web
rhachisa. the axis of an inflorescence when somewhat elongated, as in a raceme b. (in a pinnately compound leaf or frond) the prolongation of the petiole along which the leaflets are disposed c. any of various axial structures
rhachisthe spinal column, also RACHIS