Answer: tapetal

AnswerCrossword Clue
tapetalany of certain membranous layers or layered coverings, as in the choroid of the eyes of certain animals
tapetallayer of cells often investing the archespore in a developing sporangium and absorbed as the spores mature
tapetalpertaining to a tapetum ((Latin) a reflective layer of the choroid in the eyes of many animals, causing them to shine in the dark)